1) The formation of mid-level mesocyclones are thought to be very-well understood. It forms as a result of updraft-tilting of horizontal vorticity, or, more specifically, sufficient amounts of storm-relative streamwise vorticity.
2) The formation of a low-level mesocyclone is not as well understood. It is thought that it forms in a similar manner to that of the mid-level mesocyclone except that the relative roles of horizontal vorticity from vertical wind shear and that from baroclinic generation of horizontal vorticity is still being debated.
3) The formation of rotation near the ground (surface) is not well-understood at all. The tilting and stretching of horizontal vorticity is not adequate for explaining rotation near the ground. The updraft that participates in the stretching and tilting also would advect vertical vorticity upward, preventing rotation near the surface. So it is thought that a downdraft is necessary to keep a circulation near the ground.
4) The formation of the tornado after rotation has been established is better understood. It results from large increases in cyclonic vertical vorticity resulting from stretching in the updraft. With enough convergence and angular momentum, a vortex forms at mid-levels first and propagates downward by drawing in air towards its lower end. The convergence associated with the drawn in air helps to amplify vertical vorticity and cylclostrophic balance is established at the lower level. This process repeats until a vortex reaches the ground. Friction at the ground also appears to play a role by generating strong radial inflow near the ground in the vortex.
Many theories have been offered as explanations for #3, though there really is no consensus at this point.
2) The formation of a low-level mesocyclone is not as well understood. It is thought that it forms in a similar manner to that of the mid-level mesocyclone except that the relative roles of horizontal vorticity from vertical wind shear and that from baroclinic generation of horizontal vorticity is still being debated.
3) The formation of rotation near the ground (surface) is not well-understood at all. The tilting and stretching of horizontal vorticity is not adequate for explaining rotation near the ground. The updraft that participates in the stretching and tilting also would advect vertical vorticity upward, preventing rotation near the surface. So it is thought that a downdraft is necessary to keep a circulation near the ground.
4) The formation of the tornado after rotation has been established is better understood. It results from large increases in cyclonic vertical vorticity resulting from stretching in the updraft. With enough convergence and angular momentum, a vortex forms at mid-levels first and propagates downward by drawing in air towards its lower end. The convergence associated with the drawn in air helps to amplify vertical vorticity and cylclostrophic balance is established at the lower level. This process repeats until a vortex reaches the ground. Friction at the ground also appears to play a role by generating strong radial inflow near the ground in the vortex.
Many theories have been offered as explanations for #3, though there really is no consensus at this point.
The above is based on a review article that Bob Davies-Jones wrote recently, which is, I believe, unpublished (unfortunately). I suppose this thread could serve as providing information and answering questions anyone might have regarding tornado and tornadogenesis research.


